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Please Help - Exch 2010 not receiving external mail

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I have Exchange 2010 (on SBS2011) 

When I send mail to user@mycustomer.com I get the follwing NDR 550 - Mailbox unavailable. The server response was: 5.7.1 Unable to relay

- All users can send mail

- I can telnet to the server i.e MX etc all ok, firewall rules ok

- I was using POP3 (inbound) and now trying to config server to be fully SMTP

- Users have mailbox with default recipient policy addr @mycustomer.com

- I have two Receive Connectors - domain.local & mycustomer.com

Is there some setting I am missing?  DNS? Permissions? 

Really pulling my hair out at this stage - please help

Thanks in advance


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